This is the fi rst revision of Artema Walckenaer, 1837, a genus consisting of large and phylogenetically interesting species. Even though Artema is not species-rich (now eight nominal species), it has suffered from poor descriptions and synonymies. Our main goal was to gather all available material and to clarify species limits. Four species are easily distinguished from other congeners: Artema atlanta Walckenaer, 1837, the type species; A. kochi Kulczyński, 1901 (revalidated); A. bunkpurugu Huber & Kwapong, 2013; and A. nephilit sp. nov. All other species are problematic for varying reasons: species limits are unclear between A. doriae Thorell, 1881 and A. transcaspica Spassky, 1934; A. magna Roewer, 1960 and A. ziaretana (Roewer, 1960) are problematic because they are based on female and juvenile types respectively and little new material is available. The material available to us suggests the existence of a few further species; however, they are not formally described, either because of small sample sizes (Artema sp. a and A. sp. b are represented by only one specimen each) or because of unclear species limits (between Artema sp. c, A. transcaspica and A. doriae).This study is the fi rst serious step towards understanding the genus. Intensive collecting effort is needed in order to fully clarify species limits.
Bibliographical noteFunding Information:
This study was supported by the Israel Taxonomy Initiative (ITI) biodiversity survey grant, and ITI and Ben-Gurion University fellowships to SA. We thank the Steinhardt Museum of Natural History for support of BAH to visit Israel. We thank the Israel National Parks Authority for issuing the collection permits (2013/40027, 2013/40085 and 2014/40313). We thank N. Berner-Aharon for editorial help, Figs 195–200. Artema sp. c, ♀♀, epigyna, ventral (195–199) and lateral (200) views. 195. India, Rajastan, hill S Pali (CAS). 196–198, 200. Pakistan, Baluchistan, Muslim Bagh (ZMMU). 199. Sudan (AMNH). Scale lines: 0.5 mm.
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- Middle East